Trump Odds On Binary Options
Proc Natl Acad Sci U Southward A. 2009 Jan 20; 106(3): 679–684.
From the Cover
Applied Mathematics, Political Sciences
Price dynamics in political prediction markets
Saikat Ray Majumder
aNorthwestern Plant on Complex Systems, Northwestern Academy, Evanston, IL 60208;
bKellogg Schoolhouse of Direction, Northwestern University, Evanston, IL 60208;
Daniel Diermeier
aNorthwestern Institute on Circuitous Systems, Northwestern Academy, Evanston, IL 60208;
bKellogg School of Management, Northwestern University, Evanston, IL 60208;
Thomas A. Rietz
cHenry B. Tippie College of Business organization, University of Iowa, Iowa Metropolis, IA 52242; and
Luís A. Nunes Amaral
aNorthwestern Institute on Complex Systems, Northwestern University, Evanston, IL 60208;
dDepartment Chemic and Biological Technology, Northwestern University, Evanston, IL 60208
- Supplementary Materials
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Supporting Data
GUID: 7C122C6C-95C0-4FFE-B549-C616D904CFBF
GUID: F93653D2-3FCF-4F8C-BFE3-7E919EA941FA
Abstract
Prediction markets, in which contract prices are used to forecast future events, are increasingly applied to various domains ranging from political contests to scientific breakthroughs. However, the dynamics of such markets are not well understood. Here, nosotros study the render dynamics of the oldest, most information-rich prediction markets, the Iowa Electronic Presidential Election "winner-takes-all" markets. As with other financial markets, nosotros find uncorrelated returns, power-police force decomposable volatility correlations, and, usually, power-law decaying distributions of returns. Nonetheless, unlike other financial markets, we find conditional diverging volatilities as the contract settlement date approaches. We advise a dynamic binary choice model that captures all features of the empirical data and can potentially provide a tool with which one may extract truthful information events from a price fourth dimension series.
Keywords: econophysics, data assemblage, wisdom of the crowd, binary model
Prediction markets trade specifically designed futures contracts with payoffs tied to upcoming events of involvement (1). A common type of prediction market contract is a binary option contract that pays off $i if an event occurs and $0 otherwise. The contract blueprint, which differentiates them from typical futures contracts, allows prices to be used equally direct forecasts of effect probabilities (2–5).
Although betting on election outcomes was quite mutual in the United States prior to the 2d World War as discussed in ref. half-dozen, emergence of modern prediction markets, with the goal of information assemblage and revelation, tin can largely be traced dorsum to the markets introduced by the Iowa Electronic Markets (IEMs) in 1988 (seven). Since so, prediction markets have been created for ballot outcomes (seven), financial results of companies (8), scientific breakthroughs (nine), incidence of infectious disease (10), geopolitical events (ix), box office takes of movies (eleven), the outcomes of sporting events (12), and hurricane landfalls (xiii, 14), among others. They have besides been proposed for topics ranging from terrorist attacks (fifteen) to future Olympic sites (xvi). Hedge Street (17) at present trades binary option contracts on gold, silver, crude oil, and foreign exchange. More significantly, the Chicago Board of Merchandise (CBOT) recently created binary options markets on the Federal Funds target rate (18), a leading indicator of the U.S. economic system.
Given their accurateness, reaction speed, and data richness (3, nineteen–23), prediction markets provide researchers with the opportunity to precisely assess how external factors shape collective beliefs about the likelihood of a given upshot. Here, we consider the paradigmatic case of U.Southward. presidential elections. We utilise the tools of fiscal time series analysis and econophysics (24–26) to investigate the price dynamics of prediction markets with the goal of developing methods to identify the truly critical events during presidential campaigns. There are numerous known empirical regularities for price dynamics in stock, foreign commutation, article spot and futures markets (27–34). At that place is likewise some research on "ordinary" options returns (35, 36) and much on the relationship between options prices and stock returns (37–39). For details, refer to refs. 37 and 38, which survey the extensive literature on empirical selection pricing inquiry, stock options, options on stock indexes and stock alphabetize futures, and options on currencies and currency futures. Even so, empirical render characteristics for binary options—which differ considerably from other financial instruments, including ordinary options contracts*—accept not yet been documented.
As a first step toward our goal, nosotros investigate the statistical backdrop of the prices in the two most active IEM presidential winner-takes-all markets. Our empirical assay of the data for the Democratic contracts in yr 2000 and Democratic and Republican contracts in 2004 reveals that the distribution of returns decays in the tail as a power law with an exponent α ≈ 2.6. However, for the Republican contracts in twelvemonth 2000 we observe that the return distribution decays as an exponential function with a characteristic disuse scale β ≈ 0.9. We conjecture that this may have resulted from the greater influence of partisan trading for this particular contract.
Our empirical assay enables us to propose and test a dynamic binary options model with conditional leap sizes and diverging volatility. We demonstrate that the model reproduces all the chief features of the price dynamics in binary option markets. The model besides suggests a criterion for identifying extraordinary price movements arising in such markets due to significant information events and thereby raises the possibility that ane may be able to identify those events that shape a political campaign.
Maturity of Prediction Markets
Prediction markets are a relatively new forecasting tool. Yet, some markets take trade volumes similar to traditional futures markets. For example, the daily number of trades in the IEM electronic markets that we written report is comparable to the number of trades for equity options for very large companies such as IBM or DELL on the New York Stock Substitution. In fact, the number of trades in the IEM Federal Funds marketplace is much college than that for the similar CBOT binary options on rate decisions by the U.S. Federal Reserve [come across supporting information (SI) Appendix for details]. Thus, although the dollar value of the contracts traded in the IEM is small, they are very active markets. Moreover, experimental economics evidence (42) and evidence from the prediction markets themselves (21) evidence that, fifty-fifty for modest monetary payoffs, active markets reveal trader information. These facts suggest that at least large prediction markets, such equally IEM markets for U.S. presidential elections, are mature plenty to warrant analysis.
Prediction markets have been remarkably successful in correctly predicting hereafter events (3, xix, 21, 22). For example, in presidential elections prediction markets routinely outperform opinion polls (21). This generalizes to other domains as well (3, 22, 23). Moreover, prediction markets speedily incorporate new information as was demonstrated in the IEM "1996 Colin Powell Nomination marketplace" (twenty) (see SI Appendix for details). Given their large trading book, reaction speed, and accuracy, IEM, therefore, provides us with the opportunity to assess how external events shape commonage beliefs about the likelihood of a given event in the context of a political campaign.
The Data
The IEMs are real coin markets open 24 hours a 24-hour interval, 7 days a week with trading through the Internet. Trading on their own accounts, traders place "bids" to buy and "asks" to sell contracts. These orders are placed into cost- and time-ordered queues. Traders may also set the expiration of the order. If no expiration is provided, the club is removed at 11:59 PM Central Standard Time (CST) the day after the order was placed in the queue. The highest bid and lowest ask are available to all traders logged into the marketplace. Besides placing an society into the queue, a trader tin also accept the best bid (enquire) to buy (sell) a contract. All viable trades are executed immediately.
The IEM records information on every trade, including whether the trade was executed at the bid or inquire and whether there were multiple individual trades associated with a unmarried order. For convenience, nosotros build equal time-interval fourth dimension series for price, number of trades, and volume in dollars, where the time interval is τ = sixty sec. We have checked that the dynamics of equal-interval time series is similar to the time series with actual trade times.
The 2000 presidential election winner-takes-all market opened on May 1, 2000 with contracts associated with the Democratic, Reform, and Republican parties; the 2004 presidential ballot winner-takes-all market opened on June 1, 2004, with contracts associated with the Democratic and Republican parties. These markets traded binary options contracts tied to the election outcome (43, 44). Each traded contract was associated with a political party and paid $1 if that political party received the majority of the two-party or three-party pop vote.
In theory, traders in prediction markets price contracts co-ordinate to their expectations, and then the prices will be a noisy proxy for the aggregate estimated probability of the associated result†; run across ref. 45 for a more detailed discussion. Thus, the toll of the contract associated with the Democratic party indicates the probability (with some uncertainty) that the party'due south nominee volition have the majority of the 2-party vote. Note, nonetheless, that there will always exist some remainder uncertainty and, hence, prices should remain bounded away from $0 or $1 until settlement. For example, in the 1996 IEM presidential winner-takes-all markets, months in advance of the election, it was forecast that Clinton would emerge as the winner. This was reflected in the prices of the Clinton contracts, which slowly approached, but never reached, $i.
Statistical Backdrop of the Returns
In the IEM presidential election markets, contracts are finer settled on election day, which is well-known in accelerate: Nov seven for 2000 and November ii for 2004. We gear up the origin of the fourth dimension axis at these settlement dates. The times in our fourth dimension serial are so indexed as
where i = 0,1,…,Due north, and τ0 ≡ 0 is the time when the contracts are settled and τ North is when the market place opens.
We define the return at fourth dimension τ i equally
where p(τ i ) is the cost of the contract at time τ i .
Although little is know about the toll dynamics in prediction markets, in that location are three well-established facts about price fluctuations in stock markets, foreign commutation markets, and commodity markets (27–34). First, returns are uncorrelated for time scales longer than a few seconds. Second, volatilities are positively correlated over long time periods. Specifically, the correlations of the volatility decay as power laws. Third, the distribution of returns is consistent with a power-law asymptotic behavior,
For stocks, foreign exchange rates, and commodity futures, the exponent α ∼ 3 (well exterior the stable Lévy regime 0 < α < 2) (24, 34), but α ∼ 2.3 for commodity spot prices (32).
Nosotros quantify the price dynamics of the Autonomous and Republican contracts for the 2000 and 2004 elections along these three dimensions. We find that the number of trades increases dramatically toward the settlement date and that the returns in the final days of the market have significantly higher volatilities; cf. Fig. 1 C. Specifically, conditional on a given price, the volatility is higher the closer the contract is to liquidation, that is, for a given p(τ i ), the volatility diverges as τ i approaches nix (see SI Appendix for details). For this reason, we separately analyze the data in yr 2000 for the final 10 days of the market (days one-10), and for each of the previous two-month periods (days 11–70, 71–130, and 131–190). To avert issues that may ascend equally information comes in on election day, we simply clarify data up to midnight the day earlier the election [equally is normally done in the prediction marketplace literature (iii)].
Trading dynamics in the 2000 presidential ballot market. (A–C) The bold line indicates the 2000 Democrat contract whereas the thin line indicates the 2000 Republican contract. Time is counted from the settlement date. (A) Toll of the contracts in USD$. (B) Volatilities, estimated every bit standard deviation of the returns calculated on nonoverlapping 12-hour windows. (C) Daily number of trades. Note how the standard difference of the returns and the number of trades both increment markedly toward the settlement engagement.
To determine whether long-range correlations exist in the returns, nosotros use detrended fluctuation analysis (46–48, 55), which works as follows. Consider a fourth dimension series 10(t i ). One integrates this time series, generating a new time serial y(t i ), which is and so divided into blocks of size northward. In each box, one performs a to the lowest degree-squares linear fit to the information (to capture any local trends at scale due north), and determines the sum F(n) of the squares of the residuals inside all the blocks of size n. This procedure is then repeated for different values of due north. If x(t i ) can be modeled every bit independent and identically distributed (i.i.d.) Gaussian variables, one finds
Exponent values > one/2 indicate positive long-range correlations, whereas smaller values point long-range anticorrelations. For returns, we find an exponent ∼0.five. For the volatilities, which nosotros ascertain here as the absolute value of the returns, nosotros observe an exponent ∼0.7 (Fig. 2 A and B), except during the first two months of the market (days 131–190), when trading was very thin and the exponent is ∼0.5. These results are consistent with the hypothesis that the returns brandish no correlations while there are positive long-range volatility correlations, like to what is plant in other financial markets.‡
Statistics of the returns for the 2000 Democratic contract. We employ the detrended fluctuation method (meet main text and SI Appendix) to quantify the correlations. (A) Autocorrelation of the returns. The data are consequent with uncorrelated returns. (B) Autocorrelation of the volatilities. We discover ability-police decaying positive correlations for the return volatilities. This implies that periods of big volatility are more likely to exist followed by periods of big volatility than by periods of depression volatility. (C) Distribution of the positive returns. The data propose the possibility of an asymptotic ability-law decay of the distribution of the returns.
Next, we judge the power-law exponent α, defined in Eq. 3, for the return distributions. Equally shown in Fig. ii C, the render distributions in days 1–10 are wider than for the previous months. Yet, we notice that if we normalize the returns with the volatilities estimated separately in each 1 of the time periods, so the normalized return distributions follow the same functional forms. Specifically, the Kolmogorov–Smirnov (KS) test fails to reject the null hypothesis that the normalized returns are fatigued from the aforementioned distribution.§
We compute the volatility for each one of the time periods as the standard deviation of returns over that time menstruation,
where T denotes one of the time periods and 〈…〉 denotes a time boilerplate over the time menses T. The normalized returns T (τ i ) in T are so defined as
Since these normalized return distributions have the same functional forms, we compute a unmarried distribution for the positive and negative returns from the different time periods. Using the Hill reckoner (54) and bootstrapping, nosotros then get α = 2.6 ± 0.2 for the 2000 and 2004 Democratic contracts and the 2004 Republican contract (Fig. 3 A and C)¶.
Asymptotic behavior of the distribution of returns for the 2000 presidential ballot market place. (A) Double-logarithm plot of the distribution of normalized 2000 Democratic contract returns for the different time periods and for both the positive and the negative tails. The ruby lines show the 95% conviction intervals. This shows that the deviations in the tails of the distributions are consistent with the expected fluctuations. (B) Log-linear plot of the distribution of normalized 2000 Republican contract returns for the different fourth dimension periods and for both the positive and the negative tails. The carmine lines show the 95% conviction intervals. This shows that the deviations in the tails of the distributions are consistent with the expected fluctuations. (C) Double-logarithm plot of the distributions of pooled normalized 2000 Democratic contract (total line) and of pooled normalized 2000 Republican contract (dashed line). A straight line in this plot indicates a power-law dependence. (D) Same information just in a log-linear plot. A straight line in this plot indicates an exponential dependence.
Surprisingly, for the 2000 Republican contract, we discover the return distribution decays at an exponential charge per unit,
where β is the characteristic disuse scale. We observe that the tails of the return distributions disuse with the rate β = 0.9 ± 0.1 (Fig. 3 B and D). The fact that the Republican contracts are not perfectly negatively correlated with the Democratic contracts can exist understood if i recalls that the market in 2000 included a Reform party winner-takes-all contract (in improver to the Autonomous and Republican contracts).
The exponential decay of the render distribution tin exist attributed to partisan trading. For a well-performance market place in which traders take no partisan beliefs, ane would wait traders to buy (sell) Democratic and Republican contracts at approximately equal rates. However, traders affiliated with a political party tend to preferentially purchase the contract of the party with which they are affiliated and to preferentially sell the contract of the other party. While the bias in those choices is relatively small for the 2000 and 2004 Democratic contracts and the 2004 Republican contract, they are stronger for the 2000 Republican contract. Relative to other contracts, more Republican traders in 2000 trade as if they truly believe the Republican candidate is going to win, and more Democrat traders merchandise every bit if they truly believe that the Republican candidate is going to lose. Thus, while those partisan Republican traders are very willing to purchase the Republican contract, the partisan Democrat traders are very willing to sell it (run across SI Appendix, Tabular array 3).
These biases accept two consequences. Start, these traders may have on substantial risk, since their portfolios will be heavily "tilted" toward i of the contracts. 2nd, partisan traders' disability to suit new data as rapidly as nonpartisan traders (49) results in their constant willingness to purchase (or sell, depending on their bias) which prevents returns with larger magnitude from occurring. Interestingly, our findings for the 2000 Republican contract mirrors unexplained findings for the Indian stock market place. Specifically, Matia et al. (l) reported an exponential decomposable probability density office of the price fluctuations when they analyzed the daily returns for the period Nov 1994 to June 2002 for the 49 largest stocks of the National Stock Substitution in India. Our analysis suggests the hypothesis that a meaning fraction of Indian traders may hold stiff biases that determine their trading strategies.∥
The existing information practice not allow a full exploration of this hypothesis at this stage. Thus, we can non give a full explanation of why the partisan effect was stronger in 2000 than in 2004. However, studies related to cognitive dissonance (52) or confirmation bias (53) would suggest that the outcome would be stronger in elections with stronger emotional zipper to the respective candidates. This strikes united states as a promising avenue for further research.
Model
Binary options liquidate at either $0 or $1. This implies a pricing discontinuity at maturity. The value of the option will jump from the current price to either $0 or $1 the instant the dubiety is resolved. Another meaning feature of binary choice contracts is that the range of possible returns depends on the electric current price. For prices close to, for example, $1, the price can increase only past a very small-scale amount, nevertheless, it can decrease by 100%. As a effect, a plausible model must incorporate conditional asymmetric upward and down jumps with increasing volatility as i approaches the settlement date.
Let T a exist the boilerplate fourth dimension between consecutive trades and t i the time at which the i th-to-final trade occurs. The median time divergence between consecutive trades for the 2000 Democratic contract was ∼60 sec and, therefore, nosotros set up T a = 60 sec. Nosotros hypothesize that the electric current value of a winner-takes-all contract, which will settle at a value of $1 or $0, evolves according to
where γ ≥ 0 and P(t i ) refers to the value at the i th-to-terminal trade. This process is a Martingale; at any point in fourth dimension 〈P(t i )〉 = P(t i ).**
Additionally, we see that it converges to the advisable value at settlement
The model also makes information technology clear what nosotros mean by conditional diverging volatility. The variance of returns from the underlying procedure is given by
This explicitly shows that, conditional on a given price, P(t i ), the volatility is expected to exist college the closer a contract is to liquidation.
Eq. 8 models the dynamics of the "true" value of the contract. The actual price, p(t i ) will, nevertheless, deviate from P(t i ) due to noise and toll information delays. To incorporate noise we model the observed price process in the following manner
where η and ɛ are Gaussian random variables. The additive noise term, η, prevents the prices $0 and $1 from becoming absorbing states of the dynamics. For η we model a Gaussian distributed variable with null mean and a very small standard departure; the results shown were obtained for a standard departure of 0.0003. Because η≠0, the price, p(t i ), deviates slightly from a martingale process.
Past price information delays we refer to the fact that traders may not have access to the nearly current price but to a price some time in the by. Since at that place is a 15- to thirty-sec time lag for the IEM trading arrangement to update information we fix the time lag in our model to 25 sec (20).
Another upshot that also needs to be taken into consideration is that big-book bids or asks that cross the opposing queue may non trade at a single price. Instead, they will "run" through the opposing queue generating a serial of prices that all move in the aforementioned direction. Nosotros treat each such event as a single merchandise.
The model as defined higher up then has the post-obit free parameters: the exponent γ and the mean (μ) and the standard deviation (σ) of the noise term ɛ. To estimate these parameters, we run across from Eq. 11 that if we set δp(t i ) = (p(t i−1) − p(t i ))/(ane − p(t i )) for positive price changes and δp(t i ) = |(p(t i−1) − p(t i ))/p(t i )| for negative changes (where |…| announce the absolute value) then we obtain,
We can guess γ from the gradient of the linear fit in Eq. 12 which tin and so be used to calculate σ from the standard deviation of the residuals.†† Nosotros estimated γ = 0.49 ± 0.01 and σ = 1.22 ± 0.02 (refer to the SI Appendix for details).
We perform Monte Carlo simulations of the model with the estimated parameter values and detect that the model generates uncorrelated returns and power-law decaying volatility correlations, in quantitative agreement with the empirical results. We also detect that the actual toll dynamics is well bounded past 90% conviction bounds as shown in Fig. 4 C and D (for a description of this method and results from the model see SI Appendix). Additionally, nosotros find that the distribution of returns decays equally a power constabulary. Using Hill figurer and bootstrapping, nosotros estimate α = ii.3 ± 0.ii, consequent with the estimate for the empirical data.
The dynamic binary option model. (A) 2 realizations of the binary option model. (B) Cost of the 2000 Autonomous contract (black line) and 90% confidence intervals for the model. (C) Returns of the 2000 Democratic contract (black line) and 90% confidence intervals for the model. Note how some returns in the data far exceed the xc% confidence intervals from the model, as may be expected given the power-law distribution of the returns.
Discussion
The remarkable agreement between model predictions and the data may suggest a reasonably good understanding of the dynamics of prediction markets. Still, in that location is ane central feature of prediction markets neglected by the model. In real prediction markets there is true information in the grade of "known unknowns," such as the outcomes of debates or "unknown unknowns," such as revelations well-nigh the candidate'southward past, arriving at the market. These real information events tin can be viewed as exogenous processes and may be characterized by larger jumps than those arising from endogenous processes. Information technology is then plausible that the identification of sharp differences between model predictions, the endogenous events, and real information, the exogenous events, could exist used as a tool to place information arrival at the marketplace. In the context of a political contest, this approach can be used to determine which entrada events take a substantial impact on the fortunes of a item candidate.
There is another possible application of our model which we believe will have a bang-up impact in the course of a political campaign. In an election, there is a predetermined date when all the uncertainties are resolved, the settlement appointment. One may, still, realize that, in a particular election year, much of the dubiousness can exist resolved earlier than the actual settlement engagement. For example, in the 1996 presidential election, it was forecast that Clinton would sally every bit the winner about 100 days prior to the actual settlement engagement. Our model can be used to estimate this appointment by which most uncertainties are settled and, every bit a result, enable the political campaigners to judiciously assign their campaign resources.
Although our focus here is on political markets, our insights utilise to binary options markets in general and thus volition be of import for traders, exchanges, regulators, policy makers, and forecasters alike. For case, our model can be used to forecast a distribution of probable cost movements and, as a result, be used past exchanges to gear up margin requirements for traders of binary options conditional on prices and time to settlement. Another interesting aspect of our report is the possible application to crashes in financial markets. The approach to settlement date is remarkably similar to the increased volatility close to a market crash. Potentially, a generalization of our model could be used to gauge the time of a crash in these markets.
Supplementary Material
Acknowledgments.
We thank the IEM team and especially Joyce Berg (Academy of Iowa Henry B. Tippie College of Concern) for providing u.s. with the opportunity and information to perform this work, and also D. South. Bates, R. D. Malmgreen, M. J. Stringer, R. Guimera, P. Mcmullen, K. Sales-Pardo, A. Salazar, and Southward. Seaver for comments and discussions.
Footnotes
The authors declare no conflict of involvement.
This commodity is a PNAS Direct Submission.
*Binary options differ from ordinary options in 3 respects: (i) the payoff structure, (ii) the fact that there is no underlying traded asset, and (iii) pricing discontinuities at settlement, as we will bear witness below.
†Small amounts of noise may ascend from the bid–ask spread, asynchronous trading, and "stale" prices, though such factors should be pocket-size in agile markets.
‡Come across SI Appendix for details of the method and the results. To make certain that those results are not due to the non-Gaussian distribution of the returns, we randomized the time social club of the returns and reevaluated the exponent values. We notice that, for the randomized time serial, the exponent values ∼0.5 for both the returns and volatilities.
§Meet SI Appendix for the P values from the KS tests. The confidence premises in Fig. 3 A and B show that the deviations in the tails are consequent with expected fluctuations.
¶Refer to the SI Appendix for description of these and related statistical methods.
∥However, in ref. 56, R. Thou. Pan and S. Sinha have analyzed loftier-frequency tick-by-tick data for the Indian stock marketplace and plant that the cumulative distribution has a tail described by the power constabulary with an exponent ∼iii contrary to the findings in ref. 50.
**Technically, P(t i ) is the gamble neutral mensurate, but should approximate the truthful probability in the absence of significant hedging need. This implies the best forecast of the next price is the current price. In fact, the current cost is the all-time forecast of the settlement value. In context, this is Fama's weak form efficiency with a zero expected render (27). A continuous arbitrage opportunity built into the IEM restricts the adventure-costless rate to cypher. Specifically, the "unit portfolio" of both (or all 3) contracts is risk gratuitous and tin always be traded for $one cash and vice versa. Cash accounts earn nix interest. Since the aggregate portfolio is also run a risk gratuitous, it earns a nothing return and, hence, the returns to aggregate risk factors are zero. That is, all assets should earn the risk-gratuitous rate, in this case, 0. Pricing contingent claims with zero aggregate run a risk at expected value results from a simple extension of refs. 40 and 41.
††Nosotros have assumed that ɛ has hateful null. Ane might, instead, assume that the hateful is not zero and attempt to estimate information technology too. In Eq. xi, however, both the μ and γ finer scale the leap sizes relative to the remaining time controlling the speed of convergence. As a consequence, they prove very difficult to identify independently without inordinate amounts of data. Preliminary analysis indicates a correspondence between the μ and γ estimates, where the speed of convergence weighs relatively more heavily on one parameter or the other. Pairs of estimates appear to explicate the information equally. Here, nosotros choose to model mean zero noise and permit γ reflect the speed of convergence. We leave further exploration of the μ – γ relationship to future inquiry.
This article contains supporting information online at www.pnas.org/cgi/content/full/0805037106/DCSupplemental.
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